My lines of research include the bryophyte flora and vegetation in Mediterranean areas, the systematics and molecular phylogeny in bryophytes, the phylogeography of mosses, especially the origin and the diversification of species, the ex-situ conservation of bryophytes and the effects of climate change on bryophytes through the study of altitudinal transects in Mediterranean mountainous areas using cosmopolitan distributed species as well as the molecular adaptation of these species to the environmental conditions in which they develop.